Dated by CD’s annotation and by the reference to Oxenden’s enclosing specimens of Ophrys arachnites (see n. 3, below).

In his letter of 15 May 1862, Oxenden asked whether there was any way in which he could assist CD; CD’s reply has not been found, but he apparently asked Oxenden for specimens of Ophrys arachnites (a synonym of Ophrys fuciflora, the late spider-orchid). In Orchids, pp. 72–3, CD reported that, while some botanists regarded O. arachnites as ‘a mere variety of the varying Bee Ophrys’ (O. apifera), his examination of specimens of O. arachnites sent him by Oxenden had led him to conclude that it was functionally very different from O. apifera; he stated: ‘until these forms can be shown to be connected by intermediate varieties, we must rank O. arachnites as a good species’. Following the publication of Orchids, CD continued to investigate the relationship between the two forms (see n. 3, below, and letter to A. G. More, 7 June 1862, letters from G. C. Oxenden, 21 June 1862 and 8 July 1862, and letter to Daniel Oliver, 24 July [1862]).

In DAR 70: 25 there is an observational note, dated 30 May 1862, that states: 2 spikes of Arachnites sent me by M^r Oxenden, kept in water (& young flowers opened) for more than week & intentionally jarred strongly & often, the [interl] pollinia did not fall out— The Shiny lateral knobs at base of Lab[ellum]. are [interl] larger than in Bee; if these be perforated *by insects, then [interl] [‘then’ del] this w^d be important difference.— A more important one, is that the stigmatic surface is more perpendicular & therefore more open with respect to base of Labellum in the Bee than in Arachnites; evidently so as [to] allow surface to be struck by [‘own’ del] *its own [interl] pollinia in the Bee. CD reported some of these findings in ‘Fertilization of orchids’, p. 145, a paper comprising revised and additional notes on the subject keyed by page number to the first edition of Orchids (see also Collected papers 2: 142).

Louisa Harriet Cochrane was married to Thomas Barnes Cochrane, eleventh earl of Dundonald (Burke’s peerage 1980).