 Joseph Dalton Hooker, from the portrait by George Richmond, 1855 Cambridge University Library
The scientific results of the Beagle voyage still dominated Darwin's working life, but he broadened his continuing investigations into the nature and origin of species. Far from being a recluse, Darwin was at the heart of British scientific society, travelling often to London and elsewhere to attend meetings and confer with colleagues, including the man who was to become his closest friend, Joseph Dalton Hooker. Down House was altered and extended to accommodate Darwin’s growing family; and, with his father’s advice, Darwin began a series of judicious financial investments to ensure a comfortable future for all those under his care.
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 Anne Elizabeth (Annie) Darwin Cambridge University Library
Charles and Emma Darwin’s eldest daughter, Annie, died at the age of ten in 1851. Emma was heavily pregnant with their fifth son, Horace, at the time and could not go with Charles when he took Annie to Malvern to consult the hydrotherapist, Dr Gully. Darwin wrote a memorial of his daughter just one week after her death. Annie's younger sister, Henrietta, recorded her own reactions in a poignant set of notes, which Emma Darwin kept.
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 Amy Richenda Darwin (née Ruck) Cambridge University Library
1876 was the year in which the Darwins became grandparents for the first time. And tragically lost their daughter-in-law, Amy, who died just days after her son's birth. All the letters from 1876 are now published in volume 24 of The Correspondence of Charles Darwin, and a new chapter to Darwin's 'Life in letters' has been added here.
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