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To Max Ernst Wichura 3 February [1865]

Summary

He has finished MEW’s work on hybrid willows [Die Bastardbefruchtung im Planzenreich (1865)] and sends his thanks. The extreme frequency of hybrid willows is new to CD, and he finds the explanation of their numbers in certain locations ingenious.

Comments on the criticism of Gärtner’s view of reversion

and the differences between MEW and Naudin.

CD now has doubts regarding his own view that hybrids are sterile from not being perfectly accommodated to their conditions of life.

Author:	Charles Robert Darwin
Addressee:	Max Ernst Wichura
Date:	3 Feb [1865]
Classmark:	Autographia (dealers) (1986)
Letter no:	DCP-LETT-4765A

Matches: 24 hits

… He has finished MEW’s work on hybrid willows [ Die Bastardbefruchtung im Planzenreich ( …
… sends his thanks. The extreme frequency of hybrid willows is new to CD, and he finds the …
… CD now has doubts regarding his own view that hybrids are sterile from not being perfectly …
… only recently finished reading your work on Hybrid Willows, & I did not wish to thank you …
… me in many ways. The extreme frequency of Hybrid Willows is quite a new fact to me & your …
… of the development of pollen-grains in hybrids. I see you doubt Gärtner’s statements about …
… wishes to make it a general law that all Hybrids quickly revert to either parent form; I …
… in the “Origin of Species,” the view that Hybrids are sterile from not being perfectly …
… March [1865] . Though Wichura concluded that hybrid willows were generally rare in nature, …
… in the margin of this discussion: ‘more Hybrids than pure parents!! ’ (see Marginalia 1: …
… Wichura also noted the rapid germination of hybrid seeds in bare spots ( Wichura 1865 , …
… and irregularities in the development of hybrid pollen-grains, see Wichura 1865 , pp. 32– …
… for pages 38 and 39, ‘pollen gets worse & worse in offspring of Hybrids inter se & in … …
… the more complex hybrid’; CD made a similar annotation in Wichura 1865 , p. 38 (see …
… enough care to prevent pollination of a hybrid offspring by a parent plant (see Gärtner …
… Naudin’s recent observations of Datura hybrids returning to their parent forms (see DAR …
… Hybridism’ (pp. 245–78); for his conclusions about hybrid sterility, see pp. 254– …
… 67. Regarding the sterility of hybrids, CD wrote: ‘it is scarcely possible that two …
… he was still deliberating about the causes of hybrid sterility, particularly in plants. …
… Regarding Wichura’s discussion of hybrid sterility, CD wrote in Origin 4th ed. , p. …
… The above view of the sterility of hybrids being caused by two different constitutions …
… so like in every respect to that of hybrids, which affects the illegitimate offspring of …
… makes this view rather doubtful. CD was comparing the sterility of hybrids, or the …
… offspring of hybrids, with the sterility of same-form unions among dimorphic and …

From W. D. Fox [1 June 1846]

Summary

Hybrid geese.

Author:	William Darwin Fox
Addressee:	Charles Robert Darwin
Date:	[1 June 1846]
Classmark:	DAR 205.7: 280
Letter no:	DCP-LETT-588

Matches: 5 hits

… Hybrid geese. …
… W. D. Fox informs me that in some other hybrids which he had seen there was considerable …
… note : ‘Subsequently Fox tells me that all his Hybrids are exactly like each other. —’ ink …
… asked Fox for information on geese hybrids. See Correspondence vol. 2, letter from W. …
… his species book, in a discussion of the hybrids of different breeds of geese, CD stated …

From A. R. Wallace 1 March 1868

Summary

Offers enclosure demonstrating that natural selection could produce sterility of hybrids.

More on Pangenesis and the inadequacy of H. Spencer’s approach.

Author:	Alfred Russel Wallace
Addressee:	Charles Robert Darwin
Date:	1 Mar 1868
Classmark:	DAR 106: B49–50, B53–5
Letter no:	DCP-LETT-5966

Matches: 22 hits

… selection could produce sterility of hybrids. More on Pangenesis and the inadequacy of H. …
… Nat. Select. could produce sterility of hybrids . If it does not convince you I shall be …
… 2nd. The pure progeny of A F . and of B F . and 3rd. The hybrid progeny of A F . B F . …
… the increased constitutional vigour of the hybrids exactly counterbalances their imperfect …
… will be in the 2nd. generation, besides these three classes, hybrids of the 2 nd . …
… Your Book’ pencil Enclosure: 6.1 We … hybrids. 6.3] reverse question marks, pencil 9.3 …
… the question of whether sterility of hybrids could be produced through natural selection, …
… views on the subject of the sterility of hybrids as a selected quality, see Correspondence …
… animals or plants that exhibited cross or hybrid sterility ( T. H. Huxley 1863b , pp. …
… faithfully | Alfred R. Wallace— Sterility of Hybrids produced by Natural Selection 1. Let …
… intercross with each other and produce hybrids which are also quite fertile inter-se , …
… forms. 5. The result is, that in this area hybrids will not increase so rapidly as before; …
… suited to the conditions of life than the hybrids, they will tend to supplant the latter …
… to cross unions ; and this will further diminish the production of hybrids. 7. …
… degree between the first hybrids and A F . and B F . …
… respectively. In succeeding generations there will be hybrids of all degrees, …
… varying between the first hybrids and the almost pure types of A F . and B F . 16. Now if …
… will become more and more hybridised, until the whole will be hybrids of various degress. …
… 17. Now this hybrid and somewhat intermediate race , cannot be so well adapted to the …
… a severe struggle for existence, the hybrids must succomb, especially as, by hypothesis, …
… derived from A S . and B S . and of hybrid forms mainly derived from A F . and B F . ; …
… where hybridism occurs unchecked, hybrids of various degrees will soon far outnumber the …

From Thomas Glover 26 October 1857

Summary

Describes his work, which demonstrates that hybrids of Cactus are fertile.

Author:	Thomas Glover
Addressee:	Charles Robert Darwin
Date:	26 Oct 1857
Classmark:	DAR 165: 58
Letter no:	DCP-LETT-2160

Matches: 12 hits

… Describes his work, which demonstrates that hybrids of Cactus are fertile. …
… Hooker for Glover’s address to inquire about hybrid crosses of species of Cereus , a large …
… genus of cactuses. Hybrid crosses between Cereus speciosissimus and C. flagelliformis are …
… nearer Gesneria? ) impregnated with Gesneria Cooperi—the hybrid is barren. I sent …
… them a plant to Kew— I have also got hybrids from Franciscii confortiflora impregnated …
… I am Yours faithfully | Thomas Glover. ‘Hybrid impregnated by Speciossissmus very fertile’ …
… doubt their being fertile. These latter hybrids have all the form of flagelliformis— There …
… worth recording, but I have obtained a hybrid from Gloxinia tubiflora (quere, is this not …
… known as Jenkinsoni which is undoubtedly a hybrid, by speciosus on speciosissimus, having …
… it in quantities, I suspect it may be a hybrid also. Many years ago, it struck me that if …
… it is sufficient to prove that the hybrids are fertile— I also impregnated speciosissimus …
… colour— I strongly suspect it to be a hybrid, as it came from a person whom I suspect, …

From Thomas Bell Salter 25 June 1855

Summary

Discusses hybrid plants he has raised, particularly hybrids between Geum urbanum and G. rivale, which are very fertile and exhibit great variability. [See Natural selection, p. 102.]

Author:	Thomas Bell Salter
Addressee:	Charles Robert Darwin
Date:	25 June 1855
Classmark:	DAR 177: 16 (fragile)
Letter no:	DCP-LETT-1703

Matches: 12 hits

… Discusses hybrid plants he …
… has raised, particularly hybrids between Geum urbanum and G. rivale , which are very …
… Bell. 1852. On the fertility of certain hybrids. Phytologist 4, pt 2: 737– 42. [Vols. 5,6] …
… a paper on the fertility of certain plant hybrids, which was first read before the Isle of …
… roseum and Geum intermedium were the hybrid offspring of E. montanum and E. tetragonum , …
… that the case was more complex than mere hybrid origin. See letter to Arthur Henfrey, 17 …
… with foreign pollen but the resulting hybrid is very fertile & has abundantly seeded ever …
… since that they take their chance so that the hybrids, now coming up may be only the 2 d . …
… plants. I think I might have obtained hybrid seeds of Linaria if I had tried the pollen of …
… His researches further showed that these hybrids were entirely fertile and could reproduce …
… any number of generations, unlike most hybrids which will not continue to reproduce beyond …
… intermedium was, in fact, a species of hybrid origin ( Salter 1852 , p. 740). CD referred …

From [?] [?]

Summary

Hybrid fish.

Author:	Unidentified
Addressee:	Charles Robert Darwin
Date:	[?]
Classmark:	DAR 205.7: 283–4 (Letters)
Letter no:	DCP-LETT-13860

Matches: 1 hit

… Hybrid fish. …

To Nature 15 December [1879]

Summary

CD has repeated a test of whether hybrids of the common and Chinese goose are fertile inter se. Reports his success, and comments on its significance for the theory of descent.

Author:	Charles Robert Darwin
Addressee:	Nature
Date:	15 Dec [1879]
Classmark:	Nature, 1 January 1880, p. 207
Letter no:	DCP-LETT-12391

Matches: 12 hits

… salicaria, as sterile as are the most sterile hybrids. C harles D arwin Down, December 15 …
… CD has repeated a test of whether hybrids of the common and Chinese goose are fertile …
… Eyton’s success in breeding together hybrids of the common and the Chinese goose, which …
… fertility of hybrids from the common and chinese goose In the “Origin of Species” I have …
… the excellent authority of Mr. Eyton, of hybrids from the common and Chinese goose (Anser …
… recorded with respect to the fertility of hybrids, for many persons feel sceptical about …
… Goodacre, who gave me a brother and sister hybrid from the same hatch. A union between …
… parents, had not Mr. Eyton reared eight hybrids from one set of eggs. My small success …
… their very close relationship. The five hybrids, grandchildren of the pure parents, were …
… extremely fine birds, and resembled in every detail their hybrid parents. It …
… superfluous to test the fertility of these hybrids with either pure species, as this had …
… the hare and rabbit. The fertility of the hybrids in the present case probably depends to …

To George Henslow 15 [June 1866]

Summary

CD believes most strongly in reversion. J. G. Kölreuter’s, K. F. v Gärtner’s, and some of Charles Naudin’s cases leave no doubt in his mind. Forgets whether Herbert gave cases but in conversation he certainly believed in it. Thinks Gärtner is right to say reversion occurs only rarely in plant hybrids which have not been cultivated. [See 5120.]

Variation

Author:	Charles Robert Darwin
Addressee:	George Henslow
Date:	15 [June 1866]
Classmark:	DAR Library: tipped into George Henslow’s copy of Variation
Letter no:	DCP-LETT-5123A

Matches: 11 hits

… of the comparative frequency of reversions in hybrids whose parent species had long been …
… to say reversion occurs only rarely in plant hybrids which have not been cultivated. [See …
… of Kölreuter & Gärtner is distinct for Hybrids, & some of Naudin’s cases leave no doubt …
… the cases of Reversion in ordinary Hybrids appeared so clear that I have not thought it …
… case does not concern you, as it does not refer to Hybrids. — I fully believe that Gärtner …
… he says that Reversion occurs only rarely in hybrids made from plants, which have not been …
… pp. 474, 582. Naudin’s claim that all hybrids inevitably undergo reversion had been based …
… plants. However, Max Ernst Wichura’s experiments on uncultivated hybrid willows had led …
… him to question whether hybrids ever reverted to their parent forms ( Wichura 1865 , p. …
… and n. 14. For CD’s interest in the graft hybrid Cytisus adami (now + Laburnocytisus …
… 9. William Herbert published many works on hybrid plants; CD had discussed Herbert 1837 …

To M. T. Masters 10 October [1876]

Summary

Discusses views of [Alexander James] Maule on potatoes.

Discusses graft-hybrids.

Author:	Charles Robert Darwin
Addressee:	Maxwell Tylden Masters
Date:	10 Oct [1876]
Classmark:	DAR 146: 347
Letter no:	DCP-LETT-10637

Matches: 6 hits

… Discusses views of [Alexander James] Maule on potatoes. Discusses graft-hybrids. …
… to draw attention to the subject of graft-hybrids, and I saw the late article by M r . …
… Thomas Meehan’s claim to have produced hybrid potatoes by grafting was reported in …
… Meehan 1876 ). In his discussion of graft hybrids in the potato in Variation 2d ed. 1: …
… n.p. ] Meehan, Thomas. 1876. Graft hybrids. Proceedings of the American Association for …
… in animals; he was trying to produce hybrid potatoes that were resistent to disease by …

From Edward Hewitt 22 December 1857

Summary

Replies to more queries about fowl hybrids.

Author:	Edward Hewitt
Addressee:	Charles Robert Darwin
Date:	22 Dec 1857
Classmark:	DAR 166: 197
Letter no:	DCP-LETT-2193

Matches: 8 hits

… Replies to more queries about fowl hybrids. …
… natural desires, my experience tells me, Hybrids like to be solitary, and the aversion on …
… more profitably employed than in breeding Hybrids; although at the time I was making these …
… in CD’s discussion of the sterility of hybrids (see Natural selection , pp. 422, 429). CD …
… lived, and in perfect horror, I eschewed Hybrids from that very day. They are recklessly …
… parents were perfect “pets. ” The Sebright Hybrid at present seems more docile, but youth …
… as in the sire. Again altho’ I have seen Hybrids of 6 or seven years standing, I never yet …
… available to do real duty in an affray, but Hybrids do not want them particularly for this …

From Friedrich Hildebrand 18 May 1868

Summary

Sends graft-hybrid notice.

Author:	Friedrich Hermann Gustav (Friedrich) Hildebrand
Addressee:	Charles Robert Darwin
Date:	18 May 1868
Classmark:	DAR 166: 208
Letter no:	DCP-LETT-6182

Matches: 4 hits

… Sends graft-hybrid notice. …
… copies of my notices on the potato-graft-hybrid and on the apples I spoke of in my last …
… Hildebrand’s comments on the potato hybrid and the apples were published in Hildebrand …
… the implanted variety. When the graft-hybrids make flowers I shall try to fertilize them …

From J. J. Weir 13 November 1873

Summary

Hybrid Motacilla.

Case of female duck leaving mate to pair with male of another species.

Author:	John Jenner Weir
Addressee:	Charles Robert Darwin
Date:	13 Nov 1873
Classmark:	DAR 88: 179–80
Letter no:	DCP-LETT-9139

Matches: 8 hits

… Hybrid Motacilla . Case of female duck leaving mate to pair with male of another species. …
… crayon, del pencil ; ‘&’ blue crayon ‘Hybrids | *p. 2 ♀ deserting a male [ del pencil ] | …
… Weir refers to John Hubert Verrall . The hybrid has not been identified. Gallus gallus …
… two birds paired and in due time produced Hybrids, one only however survived, a male, till …
… shall trouble you with is that of a hybrid between a common male fowl, Gallus domesticus, …
… Tufted Ducks the result last year was several Hybrids between the two species, which I …
… saw and entertain no doubt of their Hybrid character, when at Lewes in the …
… Spring I saw one of these hybrid Ducks sitting and it appeared she had paired with one of …

From Peter Henderson 15 November 1876

Summary

Reports graft-hybrids in Cytisus.

Author:	Peter Henderson
Addressee:	Charles Robert Darwin
Date:	15 Nov 1876
Classmark:	DAR 166: 140
Letter no:	DCP-LETT-10673

Matches: 6 hits

… Reports graft-hybrids in Cytisus . …
… your views on the subject of “Graft Hybrids”, in reading your remarks on the subject …
… Adami, which is claimed to be a “graft hybrid”, between C. Laburnum and purpurea . This …
… a bud variation, instead of a so-called “graft hybrid”— I may state that Mr Davidson the …
… Henderson discussed graft hybrids in his Gardening for pleasure ( Henderson [1875] , pp. …
… record in favor of the theory of “graft hybrids”, but unfortuneately for that beleif, but …

Christiansen, Ditte G. and Reyer, Heinz-Ulrich. 2009. From clonal to sexual hybrids: genetic recombination via triploids in all-hybrid populations of water frogs. Evolution 63: 1754–68.

Matches: 2 hits

… G. and Reyer, Heinz-Ulrich. 2009. From clonal to sexual hybrids: genetic recombination via …
… triploids in all-hybrid populations of water frogs. Evolution 63: 1754–68. electronic …

From Edward Blyth 23 February 1856

Summary

Opposition to EB within the Asiatic Society.

Possibility of establishment of a zoological garden at Calcutta.

Has seen Gallus varius alive for the first time.

Will procure domestic pigeons for CD; could CD pay for them by returning hardy creatures, such as macaws and marmosets, which EB can sell for a high price in India?

Does not recall his authority for genealogy of the asses of Oman. If a genuine wild ass exists EB believes it will be in south Arabia.

Infertility of Irish and Devon red deer.

Details of an unusual species of wild dog.

Fertility of canine hybrids. General tendency toward hybrid sterility.

Has skins of hybrid Coracias and the parent species.

Wide-ranging species; skua found in Europe and Australia, but not in the tropics.

Author:	Edward Blyth
Addressee:	Charles Robert Darwin
Date:	23 Feb 1856
Classmark:	DAR 98: A128–A132
Letter no:	DCP-LETT-1832

Matches: 17 hits

… Devon red deer. Details of an unusual species of wild dog. Fertility of canine hybrids. …
… General tendency toward hybrid sterility. …
… Has skins of hybrid Coracias and the parent species. Wide-ranging species; skua found in …
… of the broad fact that the tendency of hybrids is to be sterile, and especially that the …
… unprolific, & has been found (in the equine hybrid) deficient of spermatozoa. And there …
… to the hybridity. Perhaps, however, other hybrids of the very same kind might be more or …
… by telling you that I have procured some hybrid skins of the Coracias , & also of the two …
… vel [ interl ] furcatus G. œnæus is a hybrid. ’ ’Zenana: ‘In India and Persia, that part …
… 1855] , on the prolificacy of the hybrids from two or three generations of intermixing …
… s later discussion of the fertility of such hybrids, see Variation 1: 234–5. In an earlier …
… me. The G. œn 〈 eus 〉 of Temminck is a hybrid between this & a common hen, often enough …
… interesting notice of experiments with canine hybrids: but the writer’s suggestion about …
… respecting the infertility of canine hybrids in the third generation. Unless you have two …
… supposing that you had a litter of Jackal-hybrids which bred inter se , it would be easy …
… of course, to ascertain if the Jackal-hybrids really presented exceptional phenomena. A …
… cultivated plants! The prolificacy of hybrids being of course liable to be similarly …
… result, & the non-fertility of some hybrids is not necessarily due to their mixed (or …

From Thomas Meehan 28 April 1880

Summary

There has been talk in American papers of CD’s admitting he was wrong about hybrid sterility. TM has presented CD’s views in the New York Independent.

Author:	Thomas Meehan
Addressee:	Charles Robert Darwin
Date:	28 Apr 1880
Classmark:	DAR 171: 113
Letter no:	DCP-LETT-12594

Matches: 8 hits

… papers of CD’s admitting he was wrong about hybrid sterility. TM has presented CD’s views …
… owning himself wrong about sterility in hybrids,—and I have taken occasion to set your …
… may also be interested in the succeeding item on hybrids in Nature. My friend Isaac C. …
… Bartram oak question—long supposed to be a hybrid—and I have asked him to send you a copy. …
… I think in some cases hybrids may be more fertile than their parents. We get here …
… item on CD’s position on the fertility of hybrids appeared in the Independent , 29 April …
… on the occurrence and fertility of natural hybrids. George Clinton Swallow was cited as …
… × soulangeana (saucer magnolia) is a hybrid of M. conspicua (a synonym of M. denudata , …

From John Henry Gurney 2 July 1856

Summary

Hybrids of Phasianus versicolor breed freely between themselves as well as with common pheasants. Has been assured that hybrids between mallards and pintails are sometimes fertile inter se.

Author:	John Henry Gurney
Addressee:	Charles Robert Darwin
Date:	2 July 1856
Classmark:	DAR 165: 259
Letter no:	DCP-LETT-1916

Matches: 6 hits

… Hybrids of Phasianus versicolor breed freely between themselves as well as with …
… common pheasants. Has been assured that hybrids between mallards and pintails are …
… selection , p. 439, CD cited two cases of a hybrid between the mallard and pintail: the …
… to me your note of the 23 d June as to the hybrids of P. Versicolor which I bought at …
… the degree of relationship between the hybrids so intermixing & by this time these degrees …
… lost sight of I am assured that the hybrids between the mallard & pintail are sometimes …

From C. V. Naudin 26 June 1862

Summary

Thanks for Orchids.

Plans to publish soon on hybrids.

Author:	Charles Victor Naudin
Addressee:	Charles Robert Darwin
Date:	26 June 1862
Classmark:	DAR 172.1: 6
Letter no:	DCP-LETT-3621

Matches: 11 hits

… Thanks for Orchids . Plans to publish soon on hybrids. …
… to his law of reversion, by which certain hybrids might become new species, but that he …
… with a similar subject, the observation of hybrids in the vegetable kingdom. I think that …
… readers about the usual termination of hybrid generations. It will be for me both a duty …
… and that, over a number of generations, the hybrid forms reverted to the parental types. …
… two specific essences in the pollen and ovules of the hybrid] ( Naudin 1865 , p. 150). …
… also stated that he had not been studying hybrids for long enough to have formed a settled …
… to his law of reversion, by which certain hybrids might become new species, but that he …
… and that, over a number of generations, the hybrid forms reverted to the parental types. …
… two specific essences in the pollen and ovules of the hybrid] ( Naudin 1865 , p. 150). …
… also stated that he had not been studying hybrids for long enough to have formed a settled …

To T. C. Eyton 2 November [1857]

Summary

Has TCE observed whether hybrids of Chinese and common forms [of geese] were wilder, or less tame, than both parents?

Author:	Charles Robert Darwin
Addressee:	Thomas Campbell Eyton
Date:	2 Nov [1857]
Classmark:	Cadbury Research Library: Special Collections, University of Birmingham (EYT/1/42)
Letter no:	DCP-LETT-2164

Matches: 5 hits

… Has TCE observed whether hybrids of Chinese and common forms [of geese] were wilder, or …
… yes” or “no”. Did you observe that your hybrids between the Chinese & common form, were at …
… that this is sometimes the case with hybrids, as with those from common & Musk Duck. Yours …
… Appendix II). Eyton had described these hybrid geese in Eyton 1837b and 1840. The question …
… Clitheroe in a letter to me states that his hybrids from the musk & common Duck “evinced a …

To George William Johnson or Robert Hogg 20 December [1862]

Summary

Thanks correspondent for sending him a strawberry hybrid.

Author:	Charles Robert Darwin
Addressee:	George William Johnson
Date:	20 Dec [1862]
Classmark:	C. H. Hughes-Johnson (private collection)
Letter no:	DCP-LETT-3870

Matches: 4 hits

… Thanks correspondent for sending him a strawberry hybrid. …
… kindness in sending me the Strawberry Hybrid, which I have planted. When the communication …
… to his having sent CD a strawberry hybrid (see n. 3, below); Johnson and Hogg were the …
… The reference is to three runners of a hybrid strawberry, which were sent to Johnson and …

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Darwin in letters, 1868: Studying sex

Summary

The quantity of Darwin’s correspondence increased dramatically in 1868 due largely to his ever-widening research on human evolution and sexual selection.Darwin’s theory of sexual selection as applied to human descent led him to investigate aspects of the…

Matches: 1 hits

… had led him to consider sterility in the offspring of hybrids to be an outcome of complex factors, …

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