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Letter 1505

Hancock, Albany to Darwin, C. R.

25 Feb 1853

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    Discusses taxonomic relations of Alcippe.

Transcription

25 Feby 1853—

My dear Sir,

I must apologize for having so long neglected to acknowledge the receipt of your long, interesting & obliging letter of the 12th inst:, but the truth is I have been more than usually engaged of late, and have only just now found the requisite leisure. I rejoice to find that notwithstanding the limited supply of specimens at your command you have succeeded in making out so many interesting points in the history of Alcippe. You have thrown much light upon the relationship of its various organs: the explation you give of the cushion on the cirrus is very satisfactory, and so is the account of the parasitical males which are certainly very curious—

Regarding the rank that this animal should hold in the Classification is undoubtedly a matter of no little difficulty, and must depend upon the view entertained of the relative value of the differet organs. The cirri are certainly characteristic or important organs in the Cirripedia, and as they are in direct relationship with the mouth, consequently with alimentary system they must bear strongly on the general œconomy— Any modification of these organs will influence the development of the parts about the mouth, as indeed is proved by Alcippe. Such [ new devpts] would therefore appear of greater importance than variations of the capitulum or peduncle, the presence or absence of which latter does not seem to be of any signification— Looking in this way on the matter I arrived at the conclusion that the [ latter] cirripedes could not be placed with either the sessile or pedunculate groups, both of which I believe nearly coincide in the characters of the cirri and mouth.

Does not Alcippe differ in a more striking manner from the various genera of the Lepadidæ than these genera differ from each other;? if so it can scarcely be placed in this family—and in truth can scarcely be called pedunculate— if placed in the order of which the Lepadidæ is a member,—the characters of the order will have to be altered.

I have said this much to explain the motive that induced me to place Alcippe apart from the two great groups of cirripedes, but my knowledge of the Class is so imperfect that I have no great confidence in my own opinion on the subject. I am therefore well pleased that this curious animal is now in such competent hands, and have no doubt that you will find for it its proper place in the Classification, how difficult soever the task—

I have not yet succeeded in preserving more specimens; but will assuredly have a supply this spring— I have spoken to two gentlemen who live on the coast and they have promised to do their best— I shall also be frequently at Cullercoats myself—
Chas Darwin Esqe

    Footnotes Add

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    f1 1505.f1
    See letters to Albany Hancock, 10 February [1853], n. 4, and 12 February [1853]. CD did not alter the description of the Thoracica but merely made an exception for Alcippe. See Living Cirripedia (1854): 563–4, where Alcippe is contrasted with Cryptophialus.
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    f2 1505.f2
    A village just north of Tynemouth, on the Northumbrian coast.
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