My dear Sir

I am infinitely obliged for your most clearly stated observations on the Bee-orchis.² It is now perfectly clear that something removes the pollen-masses far more with you than in this neighbourhood. But I am utterly puzzled about the footstalk being so often cut through.³ I sh^d suspect snails.—

I yesterday found 39 flowers & of them only 1 pollen-mass in 3 flowers had been removed, & as these were extremely much withered flowers I am not quite sure of the truth of this.— The wind again is new element of doubt.— Your observations will aid me extremely in coming to some conclusion. I hope in a day or two to receive some day-moths, on the probosces of which I am assured the pollen-masses of the Bee-orchis still adhere.—

Did you ever read D^r. Hookers paper in Philosoph. Transactions 1854 p. 259 on the curious structure of the Rostellum in Listera.⁴ D^r. H. is considerably mistaken about the use of the parts, but his facts are accurate; & it is following out the remarkable modification in structure & function of the rostellum, that I am so anxious to examine Spiranthes & Epipactis & indeed all orchids. The paper is really worth reading, though the facts in several respects are far more curious than Hooker suspected.— I wrote yesterday to thank you for the Epipactis.⁵ For the chance of your liking to look at what I have found; take a recently opened flower, drag gently up the stigmatic surface, almost any object—(the side of a hooked needle) & you will find the cap of the hemispherical rostellum comes off with a touch, & being viscid on under surface, clings to needle, & as pollen-masses are already attached to the back of the rostellum, the needle drags out much pollen. But to do this, the curiously projecting, & fleshy summits of anther-cases must at same time be pushed back slightly. Now when an insect head gets into the flower, when the flap of the labellum has closed by its elasticity, the insect would naturally creep out by the back-side of flower. And mark when the insect flies to another flower with the pollen-masses adhering to it, if the flap of labellum did not easily open & allow free ingress to the insect, it would surely rub off the pollen on the upper petals & so not leave it on stigma.— It is to know whether I have rightly interpreted the structure of this whole flower that I am so curious to see how insects act.— Small insects I daresay would crawl in & out & do nothing.—

I hope that I shall not have wearied you with these details.

With my cordial thanks | Believe me | My dear Sir | Yours truly obliged | Charles Darwin

If you would like to see a pretty & curious little sight—look to Orchis pyramidalis & you will see that the sticky glands are congenitally united into a saddle-shaped organ.— Remove this under microscope by pincers applied to footstalk of pollen-mass & look quickly at the spontaneous movement of the saddle-shaped organs, & see how beautifully adapted to seize proboscis of moth.—