Zoologist. Studied natural sciences at Oxford under George Rolleston, 1866–8; physiology at Leipzig and Vienna; morphology under Ernst Haeckel at Jena; marine zoology with Anton Dohrn in Naples, 1871–2. Fellow and tutor, Exeter College, Oxford, 1872–5; professor of zoology, University College, London, 1875–91; Linacre Professor of comparative anatomy, Oxford, 1891–8; director of the natural history departments and keeper of zoology, British Museum, 1898–1907. Knighted, 1907. FRS 1875.
Sources: ODNB. (See the bibliography for full references to sources)