- the President of the American Council of Learned Societies
- the Cambridge University Librarian
- the Head of the Cambridge University Department of History and Philosophy of Science, and
- a representative of the History of Science Society.
The Project is also served by an Advisory Committee whose members include academics, descendants of Darwin, and other supporters.
The Project’s main offices are in Cambridge University Library; since 2009 an associated group of researchers has been based at Harvard.