Physician and naturalist. Took holy orders in 1853. MD 1861. Lecturer on physiology at St George’s Hospital, 1858–69; assistant physician, 1869–72. Medical officer for health for East Hertfordshire, 1873–9. Superintendent of statistics, General Register Office, 1880–1903. Translated Aristotle’s On the parts of animals into English in 1882. Published on flower structure and mechanisms for fertilisation.